The perennial grasses can be classified as either C3 or C4 plants. These terms refer to the different pathways that plants use to capture carbon dioxide during photosynthesis. All species have the more primitive C3 pathway, but the additional C4 pathway evolved in species in the wet and dry tropics. The first product of carbon fixation in C3 plants involves a 3-carbon molecule, whilst C4 plants initially produce a 4-carbon molecule that then enters the C3 cycle. Why are these differences important?
These differences are important because the two pathways are also associated with different growth requirements. C3 plants are adapted to cool season establishment and growth in either wet or dry environments. On the other hand, C4 plants are more adapted to warm or hot seasonal conditions under moist or dry environments. A feature of C3 grasses is their greater tolerance of frost compared to C4 grasses. C3 species also tend to generate less bulk than C4 species; however, feed quality is often higher than C4 grasses. Differences between C3 and C4 species are shown in Table 1.
C3 | C4 | |
---|---|---|
Initial molecule formed during photosynthesis | 3 carbon | 4 carbon |
Growth period | Cool season or yearlong | Warm season |
Light requirements | Lower | Higher |
Temperature requirements | Lower | Higher |
Moisture requirements | Higher | Lower |
Frost tolerance | Higher | Lower |
Feed quality | Higher | Lower |
Production | Lower | Higher |
Examples | weeping grass and common wheatgrass | kangaroo grass, red grass and wire grass |
The presence of both C3 and C4 species can be desirable in a pasture as they can occupy different niches (e.g. C3 species are often more abundant in the shade of trees and on southerly aspects, while C4 species often dominate full-sun conditions and northerly aspects) and thereby provide greater groundcover across a range of conditions. It is not uncommon to find both C3 and C4 species in one paddock. This has advantages in providing a broader spread of production throughout the year for both grazing enterprises and native animals.