Paul Greenwood

Research interests

  • Productivity and efficiency at pasture
  • Consequences of fetal and neonatal development  including developmental programming
  • Influences on growth, body and carcass composition, meat quality and skeletal muscle development
  • Biology underpinning gene markers
  • Livestock behaviour
  • Livestock phenomics


Paul Greenwood is a Senior Principal Research Scientist in the Extensive Livestock Industries Unit within NSW DPI Livestock Systems, and an Adjunct Professor in the School of Environment and Rural Science at the University of New England.

Dr Greenwood has a joint appointment with CSIRO Agriculture and Food in Armidale where he leads a major new research initiative on efficiency at pasture. This project area is focussed on the use of wireless sensor networks to estimate intake of pasture from behaviours, and enhanced productivity and efficiency of grazing livestock and grazing systems. This work also contributes to development of livestock phenomics facilities in commercial grazing environments to support the use of genetic and genomic enhanced selection programs, and improvements in the precision of livestock and pasture management including input data for prediction and simulation models and revisions to Australian Feeding Standards. This program of work has recently resulted in Meat & Livestock Australia Donor Company funding of $3.3 million to Dr Greenwood for NSW DPI, as part of a collaborative $7.4 million project with CSIRO on Improving Profit from Pasture through Increased Feed Efficiency, which will run until 2023 within the NSW DPI-CSIRO-UNE Livestock Productivity Partnership.

Dr Greenwood has headed NSW DPIs research efforts on “Biological validation and gene expression underpinning gene discovery for beef yield and quality characteristics” and “Regulation of growth, carcass composition and beef quality” within the Australian Beef Cooperative Research Centres (Beef CRCs). His research team has undertaken extensive work on long-term consequences of fetal and neonatal growth and nutrition in ruminants, and he has contributed to research programs on temperament and stress in relation to productivity, efficiency and meat quality. His team has also undertaken research on cellular development of muscle in meat producing species.

Dr Greenwood was a co-recipient of an Australian Research Council Eureka Prize for Research by an Interdisciplinary Team, within the Meat Standards Australia (MSA) Pathways Team for his gene marker and developmental studies within the Beef CRCs. He has presented the McClymont Lecture of the Australian Society of Animal Production, and was a member of the ARC/NH&MRC Research Network on Genes and Environment in Development and the International College of Gravida. He is regularly invited to present his research at national and international conferences and symposia.


  • PhD (Cornell University)
  • MScAgr (University of Sydney)
  • BScAgr (University of Sydney)

Current projects

  • Productivity and efficiency at pasture: Development and application of sensor technologies and livestock phenomics facilities for enhanced productivity and efficiency at pasture
  • Globally competitive Australian meat value chains enabled by advanced measurement technologies and integrated feedback systems
  • Biological validation and gene expression underpinning gene discovery for beef yield and quality characteristics, including biomarker identification
  • Genetic, nutritional and other impacts on growth, body and carcass composition, meat quality and muscle development  in meat producing species

Recent publications

Gotoh T, Greenwood PL, Baik M (2018) Editors. Global Prospects for Beef Production. Special Issue of the Asian-Australasian Journal of Animal Sciences31(7)

Greenwood PL, Gardner GE, Ferguson DM (2018) Current situation and future prospects for the Australian beef industry – A review. Asian-Australasian Journal of Animal Sciences31:992-1006 [invited review]

McGavin SL, Bishop-Hurley GJ, Charmley E, Greenwood PL, Callaghan M (2018) The effect of GPS sample interval and paddock size on estimates of distance travelled by grazing cattle in rangeland Australia. The Rangeland Journal 40:55-64

Rahman A, Smith DV, Little B, Ingham AB, Greenwood PL, Bishop-Hurley GJ. (2018) Cattle behaviour classification form collar, halter, and ear tag sensors. Information Processing in Agriculture 5:124-133

González-García E, De Oliveira Golini V, Hassoun P, Bocquier F, Hazard D, González LA, Ingham AB, Bishop-Hurley GJ,Greenwood PL (2018) An assessment of walk-over-weighing to estimate short-term individual forage intake in sheep. Animal 12:1174-1181

Greenwood PL, Paull DR, McNally J, Kalinowski T, Ebert D, Little B, Smith DV, Rahman A, Valencia P, Ingham AB, Bishop-Hurley GJ (2017) Use of sensor-determined behaviours to develop algorithms for pasture intake by individual grazing cattle. Crop & Pasture Science 68:1091-1099

Greenwood P, Clayton E, Bell A (2017) Developmental programming and beef production. Animal Frontiers 7:38-47 [invited review]

Greenwood PL, Bishop-Hurley GJ, Gonzalez LA and Ingham AB (2016) Development and application of a livestock phenomics platform to enhance productivity and efficiency at pasture. Animal Production Science56:1299-1311 [invited review]

Bell AW and Greenwood PL (2016) Prenatal origins of postnatal variation in growth, development and productivity of ruminants.Animal Production Science56:1217-1232 [invited review]

Greenwood PL, Siddell J, Walmsley BJ, Geesink GH, Pethick DW, McPhee MJ (2015) Post-weaning substitution of grazed forage with a high-energy concentrate has variable long-term effects on subcutaneous fat depths and marbling in Bos taurus genotypes. Journal of Animal Science 93:4132-4143

Greenwood PL, Valencia P, Overs L, Paull DP and Purvis IW (2014) New ways of measuring intake, efficiency and behaviour of grazing livestock. Animal Production Science54:1796-1804

Guo B, Kongsuwan K, Greenwood PL, Zhou G, Zhang W and Dalrymple BP. (2014) A gene expression estimator of intramuscular fat percentage for use in both cattle and sheep. Journal of Animal Science and Biotechnology 5:35

Nattrass GS, Cafe LM, McIntyreBL, Gardner GE, McGilchrist P, Robinson DL, Wang YH, Pethick DW and Greenwood PL. (2014) A post-transcriptional mechanism regulates calpastatin expression in bovine skeletal muscle. Journal of Animal Science 92:443-455

Bell AW and Greenwood PL (2013) Optimising maternal cow, grower and finisher performance in beef production systems. pp. 51-72. In: Optimisation of Feed Use Efficiency in Ruminant Production Systems. Makkar HPS and Beever D (Editors). FAO Symposium 27 November 2012, Bangkok, Thailand. FAO Animal Production and Health Proceedings, No. 16. FAO, Rome and Asian-Australasian Association of Animal Production Sciences [Invited Review]

Greenwood PL, Cafe LM, McIntyre BM, Geesink GH, Thompson JM, Polkinghorn R, Pethick DW and RobinsonDL (2013) Molecular value predictions: associations with beef quality, carcass, production, behavior, and efficiency phenotypes in Brahman cattle. Journal of Animal Science 91:5912-5925

Robinson DL, Cafe LM and Greenwood PL (2013) Developmental programming in cattle: Consequences for growth, efficiency, carcass, muscle and beef quality characteristics. Journal of Animal Science 91:1428-1442 [Invited Review]

Robinson DL, Cafe LM, McIntyre BM, Geesink GH, Barendse W, Pethick DW, Thompson JM, Polkinghorn R and Greenwood PL (2012) Production and processing studies on calpain-system gene markers for tenderness in cattle: Taste panel assessments of meat quality. Journal of Animal Science 90:2850-2860

Professional associations and activities

  • Australian Society of Animal Production
  • American Society of Animal Science
  • European Association of Animal Production
  • The Grassland Society of New South Wales

Fields of research

  • 070201 Animal Breeding
  • 070204 Animal Nutrition
  • 070203 Animal Management
  • 070202 Animal Growth and Development
  • 070702 Veterinary Anatomy and Physiology

Australian and New Zealand Standard Research Classification (ANZSRC)

Keyword/phrase list of research interests

  • Efficiency at pasture
  • Fetal and neonatal growth and development
  • Developmental programming
  • Fetal programming
  • Muscle biology
  • Meat quality
  • Body composition
  • Behaviour
  • Gene markers and genomics
  • Livestock phenomics

Contact details

Phone: (02) 6776 1374
Fax: (02) 6770 1830 
Mobile: 0438 645 349
Location: Armidale Livestock Industries Centre,  JSF Barker Building,  Trevenna Road,  University of New England